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It begins with a slow introduction that foreshadows the themes of the main first movement. The sonata opens with quiet intensity, its serious yet rhythmically energetic theme building up to a transition passage that all but explodes with passion. Then the clarinet introduces a second theme group over an accompaniment based on the first theme. After some anguished passagework, the exposition concludes with a gorgeous, singing codetta theme. Each of the above ideas gets a fair share of the development section, in which Mendelssohn shows his contrapuntal chops. A poignant cello statement concludes the development and overlaps into the recap, where the explosive transition is omitted. The coda begins like a repeat of the development, then accelerates to the thrilling, closing chords...but wait! The music hasn't stopped!
Yes, folks, Mendelssohn drops a BOMBSHELL! Instead of bring the first movement to a solid ending, he makes a transition back to the slow introduction - lets it die away - then, without allowing a break between movements, he goes straight into the scherzo! So the intro to the first movement also serves as a bridge to the second. You see how craftily old Felix unifies his material?
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Movement III is an über-Romantic slow movement. Its first subject is a warm, long-drawn-out, almost gushy melody that seems likely to spin endlessly out of a couple of brief opening phrases. A second idea, beginning with a horn call, has a more stately aspect, and at times picks up a tragic tone. Both themes are drawn together by the same gentle, sympathizing sort of musical "paragraph ending" - what music snobs stuffily refer to as "cadential extension" - material so memorable that it almost sounds like a theme, but that unmistakably serves to delay the period at the end of the sentence.
Movement IV is one of those movements that is supposed to have oodles of themes - somewhere I read the figure "at least four" - but it hits me as having the usual two themes, plus some dramatic transition material. The first theme group, however, has two distinct ideas in it: the opening gesture with its teasing character, and the clucking idea that grows out of it. The second theme, introduced by the oboe, turns out to be more important. The first time it comes around, it is grouped with a truly rockin' trumpet reply. The development section is eerily quiet to begin with, but it builds and builds as the second theme assumes increasing significance. It finally works its way to what sounds like a musical decision; then immediately goes all slow and quiet, dying out after a truly strange echo of the second theme. Then, Mendelssohn drops his second BOMBSHELL by closing the symphony with a coda based on a seemingly brand-new, chorale-like theme. This noble tune does, in fact, have some similarity to material heard earlier in the symphony - yet another shrewd experiment in pulling the symphony's movements together in an artistic unity.
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EDIT: Here is Kurt Masur conducting the finale of this symphony with the Leipzig Gewandhaus Orchestra:
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